Celestia and Luna are natural born alicorns, and from that comment Celestia makes about Flurry Heart, it's just because there have been no alicorns born in Equestria, since the sisters were born before Equestria was created. Celestia and Luna were born and raised with alicorn knowledge, such as the royal voice and the knowledge that they age slower than the regular pony. They also had an abundance of other knowledge, learning about the tree of harmony before they saw it for themselves. Somehow, other alicorns had disappeared, though it doesn't seem to be anything traumatic as their writings don't indicate any sort of thing. Celestia and Luna spent their younger years in Canterlot amongst themselves, playing pretend as royals and making it by themselves, perhaps stumbling across other ponies and creatures. Word makes it to the newly forming Equestrian nation, especially to the ears or Starswirl the bearded. Only those well versed had heard of alicorns before, and hearing that there were two young alicorns among Equestria, Starswirl finds them to be perfect candidates for the new rulers of Equestria. Along with being able to study and research true alicorns up close. Our story aligns with cannon, growing up, earning their cutie marks and taking the reigns of the celestial bodies, defeating
Now I like to follow even the smallest scraps of cannon… until there's some idea I like following better. Luna and Celestia are in all ways, the dark and light. Luna's inconsistences with her cutie mark can be chocked up to her being being fully intertwined with darkness. Celestia and Luna's unique abilities come from this, and effect their appearance. But I also just reeeally like Luna being part bat-pony, or at least sharing some characteristics. This comes from her transformation into Nightmare, fully becoming darkness. Also makes sense with the fact she can communicate with animals, or at least feels fitting.
In terms of what I changed about their designs, I added juuuuust a bit. I reordered Celestia's mane colors to look a little more pretty, along with adding a yellow gradient cause it looks pretty, but also represents her age. I gave Celestia markings to mirror Luna's too. And speaking of markings, something you can see here is them without their embellishments. I wanted to draw them here since I changed them up a bit, especially on Luna. They have what are called The Element's Scars. Celestia's is more severe than Luna's due to her taking control of all the elements to banish Luna. Their necklaces are tailored to cover these specific markings, especially Celestia's.
And there's my ramblings! Sorry for the absence, I've been dealing with some shit and school